MEKARN MSc 2001-2003

Citation of this paper

Evaluation of feed selection on nutrient intake, growth rate and egg performance of confined Tau Vang and Tam Hoang chickens

Evaluation of feed selection on nutrient intake, growth rate and egg performance of confined Tau Vang and Tam Hoang chickens

Nguyen Thi Thuy 1 and Brian Ogle2

1College of Agriculture, CanTho University, CanTho, Vietnam
nthithuycn@ctu.edu.vn
2
Department of Animal Nutrition and Management, Swedish University of Agricultural Sciences, Uppsala, Sweden.


Abstract

Two feeding experiments were conducted at the experimental farm of the Agricultural Faculty of Can Tho University in the South of Vietnam, to determine the effect of feed selection on the nutrient intake, growth rate and egg performance of confined improved (Tam Hoang) and local (Tau Vang) chickens.

Experiment 1 (growing period) was conducted with 166 Tam Hoang and 160 Tau Vang female chickens from 5 to 18 weeks of age. The birds were allocated at random to 2 treatments in a 2 * 2 factorial experiment. The first factor was breed (Tam Hoang and Tau Vang), and the second factor diet, including: "Mixed Diet", consisting of broken rice, fish meal and oil extracted soya bean, with oyster meal and bone meal; "Separate Diet", including the same feedstuffs supplied separately in 3 feeders, and with oyster and bone meal supplied together with the broken rice. There were thus 4 treatments with 4 replications, and 20 birds per experimental unit (pen). The diets were supplied ad - libitum from 5 to 18 weeks of age when 1 bird/pen was slaughtered for carcass evaluation.

Total DM and metabolisable energy (ME) intakes were not significantly different between breeds and treatments. Crude protein and lysine, methionine, leucine and threonine intakes were higher for the mixed diet than for the separate diet. The chickens on the separate diet ate more broken rice, and less soya bean and fish meal compared to the mixed diet. The ratio of CP to ME intake was not different between breeds, but was significantly different between diets. The ratios were 12.7 and 13.7 g CP/MJ ME (P>0.05) for the Tam Hoang and Tau Vang breeds, respectively, and 14.8 and 11.6 g CP/MJ ME (P<0.001) for the mixed and separate diets, respectively.

The average daily gain for the Tam Hoang was significantly higher than for the Tau Vang (20.5 g compared with 10.6 g/day, respectively). However, there was no significant difference (P>0.05) between the mixed and separate diets (15.9 g compared with 15. 3 g/day, respectively). The feed conversion ratios (FCR) were significantly different between treatments, at 3.2 and 4.1 kg feed/kg gain (P<0.05) for the mixed and separate diet, respectively, but were not significantly different between breeds (2.95 and 3.60 kg feed/kg gain (P>0.05) for the Tam Hoang and Tau Vang, respectively).

The Crude protein intake from separate diet was lower than that from the mixed diet, as a result the FCR from separate was higher than that on the mixed diet. However, the average daily gain was similar.

Experiment 2 was carried out after the growing period, from 18 weeks of age onwards, with 72 improved (Tam Hoang) and 75 local (Tau Vang) laying hens by randomly allocating surviving birds, which continued on the same dietary treatments as in the growing period. The dietary treatments were the same as for the growing period, except that maize replaced broken rice to improve yolk color. There were 10 birds per pen (9 hens and 1 cockerel) and the total pen number was 16. Data were collected for 20 weeks of the laying period.

Daily intakes of DM, CP and ME were 15 - 17 % higher for the improved Tam Hoang hens compared with the local hens. The hen - day production and egg weight for the Tam Hoang were higher than for the Tau Vang, 32.7 % and 44.5 g compared with 24.9 %and 39.5 g, respectively (P<0.05). However, there were no significant differences between the mixed and separate diets (29.7 and 42.3 g compared with 27.9 % and 41.7 g, respectively).

The feed consumption and feed cost/kg eggs were not significantly different between treatments and breeds. The feed cost per kg eggs for the Tam Hoang was 15,302 VND, and for the Tau Vang 17,665 VND/kg eggs.

Although the DM and CP intakes of the hens from separate diet were lower than that from mixed diet, but the egg performance were similar. Tam Hoang has higher egg performance than Tau Vang, although DM, CP and FC/kg egg were non significant differences.

Key words: Improved breed, local breed, separate feeds, mixed diet, growth rate, hen - day production, laying hens.

1. Introduction

Nutrition is of major importance in raising chickens, as feed accounts for 60 - 90 % of the total production costs (Gunaratne et al., 1992). However, it is difficult for farmers to formulate diets that match the birds' requirements and production purpose, as it requires knowledge of requirements as well as of the nutrient composition of the feed ingredients. Free choice feeding is a much more natural and simple system of feeding, since each bird can fairly accurately select its intake and balance of nutrients to meet its particular physiological requirements, and free choice feeding also offers considerable economic advantages to the farmer (Cumming et al, 1987) because the dietary ingredients offered can be simple, for example broken rice, paddy rice and soya beans, which are usually cheap and available on small farms and can be fed separately for the chickens to select. Also of course the farmer does not need to formulate the diets himself if the birds are allowed to select from different feedstuffs.

Feed supplementation is a major problem in rural poultry production and it has been calculated that scavenging birds are usually able to find feed for their maintenance, but that higher levels of production require supplementation (Dessie ,1996). The ability of the chicken to select a balanced diet from different foods has been demonstrated in broiler chickens and adult laying hens, but some of these studies have given varying results, particularly between different breeds. According to Hong Samnang (1997), for example, local breed chickens ate less protein supplement and more broken rice than improved breeds, while exotic breeds consumed more protein. However, there is little information on this subject and few studies comparing the feed selection and performance of local and improved chickens. Therefore, this experiment, comparing the effects of choice feeding on the growth rate and egg performance of local Tau Vang and improved Tam Hoang breeds, was carried out in on- station conditions in order to:

2. Materials and methods

2.1 Experimental design and birds

Two on-station experiments were conducted at the experimental farm of Cantho University. Experiment 1 was carried out in the growing period (5 to 18 weeks of age) and Experiment 2 in the laying period, from 19 to 39 weeks of age.

Experiment 1 was carried out using two breeds of chicken, the Tau Vang (local) and Tam Hoang (improved). Up to the age of 4 weeks, the chickens were raised on commercial farms and given a concentrate feed ad - libitum and vaccinated against common diseases such as Gumboro, Newcastle disease and Fowl pox. In total 324 female and 30 male chickens of the two breeds at 4 weeks of age were chosen and brought to the experimental farm. They were de-beaked before the start of the experiment and were fed a 50:50 mixture of the commercial and experimental diets for one adaptation week. The chickens were weighed initially and data were collected from five weeks of age. Antibiotics were also supplied in the water to prevent common diseases.

The experimental design was factorial with 2 factors: the first one was breed, including a local (Tau Vang) and an improved (Tam Hoang) breed. The second factor was diet, including the same feed ingredients, either mixed or provided separately. The ingredients were broken rice, fish meal and oil extracted soya bean meal. There were thus four treatment groups with 4 replications, giving 16 pens with 20 birds / pen (experimental unit). The experiment was a completely randomized design (CRD).

Experiment 2 was carried out using around half of the surviving chickens from Experiment1, and continued from the end of the growing period. It was also carried out on the experimental farm of Cantho University, from July to December 2002. In Experiment 2, the same birds and treatments from the growing period were used, except that broken rice was replaced by maize meal. In total 75 hens / breed were kept in the same pens (9 hens /pen) as in Experiment 1. The formulation of the mixed diet was changed to meet the nutrient requirements of laying hens.

2.2 Experimental diets and management

In the preliminary period (from day 1 to 28 after hatching) the chickens were fed a commercial starter diet ad-libitum, which contained 18 % CP and 12 MJ/kg DM. The chickens were transferred to the experimental farm at 28 days of age and given the experimental diet for 1 week of adaptation.

The treatments were:

The mixed diet in the growing period contained 17.5% CP and 12.9 MJ/kg.

The composition of the mixed diet in the laying period was: Maize meal (70.8%), fish meal (12.0 %), Soya bean meal (10.0 %), bone meal (2%), oyster meal (5%) and premix (0.2%). This formula was calculated to provide 17.6 %CP, and 12.9 MJ/kg, which are the requirements of laying hens according to NIAH (1995). Both treatments included a vitamin premix.

2.3 Housing and environment

The chickens were confined in pens constructed from trellis and cast nets and bamboo, with concrete floors covered with about 20 cm of rice husks for bedding, with an average density of 4 chickens/ m2 for the growing period and 2 birds per m2 for the laying period. Natural light was used in the daytime and artificial light (electric bulbs) at night, with an intensity of 3 W/m2 at floor level during the whole experimental period. The temperature inside the house varied between 25 - 37 0C. At the time of Expt 1 the weather was very hot, especially in the middle of the day, and this resulted in some signs of heat stress, especially in the Tam Hoang. Water was supplied ad - libitum in plastic tube drinkers. Feeders used were round plastic basins 35 cm in diameter and 40 cm high, and both feeders and drinkers were cleaned and refilled daily in the morning.

2.4. Diets and feeding

The chemical composition and nutrient values of ingredients and diets are shown in Table1. Mixture 1 was used for the period from 5 weeks up to 18 weeks of age, and mixture 2 was used from 18 weeks of age and throughout the laying period.

2.5. Feed sample analyses and calculations

The mixed diet and feedstuffs for both experiments were analyzed for dry matter (DM), crude protein (N*6.25), crude fibre (CF), ether extract (EE), and ash, Ca, and P by standard AOAC methods (AOAC, 1990). Analyses of neutral detergent fibre (NDF) of the mixed feed and ingredients were also done, following the procedure of Goering and Van Soest (1991). Representative samples of the mixed diet, broken rice, oil extracted soya bean and fish meal were sent to the Animal Nutrition Department of the National Institute of Animal Husbandry in Ha Noi for analysis of amino acids by using a method of high performance liquid chromatography (HPLC).(Table 2).

Metabolizable energy (ME) contents of the diets were calculated from chemical analysis data using the equation of Janssen (1989): ME (Kcal/kg)

Broken rice ME = 46.7 x DM- 46.7xAsh -69.55 x CP+ 42.95 x EE-1.95 x CF

Soya bean ME = 37.5 x CP + 70.52 x EE + 14.9 x NFE

Fish meal ME = 35.87 x DM - 34.08 x Ash + 42.09 x EE.

2.6. Feed intake and growth measurements

All 20 chickens in each pen were weighed together as an experimental unit weekly from 5 weeks of age up to first egg. Daily feed intake was calculated according to the total feed consumption of all the chickens in each pen. Fresh feed was offered every day and feed refusals were recorded every week at the same time as the birds were weighed, and the feed consumption calculated. At the end of Expt 1, one chicken in each pen with an average live weight was slaughtered for evaluation of the carcass and internal organs. From 19 weeks of age, treatments were the same as for the growing period, except that maize replaced broken rice in both diets. All eggs from 19 to 38 weeks of age were collected daily from every pen and weighed to calculate the feed consumption per kg of eggs, and mean egg weight.

2.7. Mortality and behavior

The initial number of birds and number remaining at the end of Expt 1 were recorded to calculate mortality, and eating behavior and incidence of feather picking and disease were also observed and recorded.

2.8. Statistical analysis

The data were subjected to analysis of Variance (ANOVA) by using the General Linear Model (GLM) and regression analysis of MINITAB.13 reference Manual Release 12(1998).

2.9. Economic analysis

Economic analyses were done by using current prices in Vietnamese Dong (VND) to calculate the differences in total income and total expenses between the two treatments and breeds (including feed, chicks, labor, vaccines and medicines) and net profits.

3. Results

3.1 Experiment 1. Growing period

3.1.1 Chemical composition of feedstuffs

The chemical compositions values of the broken rice, oil extracted soya bean, fish meal and mixed feed are shown in Table 1. Broken rice, soya bean, fish meal and mixed feeds had average dry matter (DM) contents of 87, 89, 89 and 87 %, respectively. The broken rice used was high in ME (14.4 MJ/kg DM), and low in fiber, lysine and methionine. Oil extracted soya bean had a fairly high crude protein content (41.0%) and was relatively high in lysine (16.5 g/kg DM) and ME. The fish meal was ground from small fish and was of fairly good quality, with a crude protein content of 48.8 %. It was bought from the coastal area and dried before grinding. It was quite high in Ca, P, lysine and methionine (5.3, 2.5, 30.0 and 14.4 g/kg respectively). Broken rice and fish meal are both used commonly in animal feeds in Vietnam.

In the laying period, maize meal replaced broken rice, as maize is also a good energy feed source and supplies carotenes for better yolk color. The chemical composition of the ingredients and the mixed diet are shown in Table 1. Maize meal, soya bean and fish meal had average dry matter (DM) contents of 85.9, 87.0, 85.7 and 86.3 %, respectively. The CP (%) and ME contents (MJ/kg) of the maize meal, soya bean and fish meal and the mixed diet were 9.0 and 13.8, 40.7 and 11.95, 48.5 and 9.15 and 17.6 and 12.85, respectively. Soya beans and maize were bought and stored in the grain form, and ground weekly immediately before mixing.

3.1.2 Feed intake

The results in Table 3 and Table 4 show that the total dry matter intakes (g/day/bird) of the Tam Hoang and Tau Vang breeds were not significantly different (P>0.05) when the initial weight of the chickens was taken into account. Also the daily DM feed intakes on the mixed and separate feeding systems were not significantly different. The total DM intakes for the Tam Hoang and Tau Vang birds, respectively, were 61.2 and 40.1 g/day, and 49.9 and 51.4 g/day for the mixed and separate diet, respectively.

Crude protein intake was not significantly different between breeds but was significantly different between diets. CP intake was 9.14 g/day and 6.33 g/day (P>0.05) for the Tam Hoang and Tau Vang, respectively, and for the mixed and separate diet was 8.51 and 6.95 g/day, respectively (P<0.001). The broken rice, fish meal and soya bean intakes were 35.9, 6.5 and 6.5 g/day for the mixed diet, and 43.4, 4.8 and 2.3 g/day, respectively, for the separate diet, and were significantly different between diets (P<0.001).

Amino acid intakes were not significantly different between breeds, but were significantly different between diets. Lysine, methionine, threonine, histidine and arginine intakes were 0.42 and 0.32, 0.21 and 0.14, 0.32 g/day and 0.24, 0.37, 0.23, 0.67 and 0.48 g/day for the Tam Hoang and Tau Vang breed, respectively. Daily intakes of lysine, methionine, threonine, histidine and arginine were 0.42 and 0.32, 0.19 and 0.16, 0.31 g/day and 0.23, 0.34 and 0.25 and 0.64 and 0.50 g/day for the mixed and separate diets, respectively (P<0.05).

The ME intakes were not significantly different between breeds and diets, and were 0.58 and 0.61 MJ/day for the mixed and separate diet, and 0.72 and 0.47 MJ/day for the Tam Hoang and Tau Vang breeds, respectively.

3.1.3 Average daily gain (ADG)

Growth rates were not significantly different between diets, and were 15.8 and 15.3 g/day (P>0.05) for the mixed and separate diets, respectively. However, there was a highly significant difference between breeds, and ADG were 20.5 and 10.6 g/day (P<0.01) for the Tam Hoang and Tau Vang, respectively.

ME/gain was not significantly different between the Tam Hoang and Tau Vang (34.9 and 43.1 MJ/kg gain, respectively (P>0.05).The ME/gain ratio for the mixed diet was 37.0 MJ/kg, while for the separate diet it was 40.9 MJ/kg gain (P<0.001).

The CP/gain ratios were not significantly different between breeds and were 430 and 590 g/kg gain for the Tam Hoang and Tau Vang, respectively. However, the CP needed for one kg gain for the separate diet was higher than for the mixed diet, and was 550 and 480 g CP/kg gain, respectively.

3.1.4 Feed conversion ratio and carcass evaluation

FCR were not significantly different between breeds, and were 2.95 and 3.61 kg feed /kg gain for the Tam Hoang and Tau Vang, respectively. However, they were significantly different between dietary treatments (P<0.05), and were 3.15 and 3.41 kg feed/kg gain for the mixed and separate diets, respectively.

The carcass percent was significantly different between diets, but not for breeds. The carcass proportions were 69 and 66 % (P<0.05) for the mixed and separate diet, respectively and 72 and 63 % (P<0.05) for the Tam Hoang and Tau Vang, respectively. The abdominal fat weights were higher (P<001) for the birds given the separate diet than the mixed diet, and were 43.1 and 85.3 g for the mixed and separate diet respectively, and 99.4 and 29.1 g for the Tam Hoang and Tau Vang breed, respectively (P<0.01).

3.1.5 Benefit analysis and mortality for the growing period

The difference in feed cost/kg gain was not significant (P>0.05) between diets, the values being 8,611 and 8,677 VND/kg gain for the mixed and separate diet, respectively. However, there was a significant difference between the Tam Hoang and Tau Vang, and the cost for 1 kg of Tam Hoang gain was 8,148 VND while for the Tau Vang it was 9,140 VND/kg gain.

The mortality was 13.4 % and 5.0 % for the Tam Hoang and Tau Vang, respectively (P<0.05). However, there were no significant differences in mortality between diets, at 9.8% and 8.6 % for the mixed and separate diet, respectively (P>0.05).

3.2 Experiment 2. Laying period

3.2.1. Feed intake

The total daily DM, CP and ME intakes were not significantly different between breeds, and were 78.2 g, 13.0 g and 1.02 MJ compared with 64.8g, 11.0g, and 0.84 MJ for the Tam Hoang and Tau Vang, respectively. The proportions of maize, soya bean and fishmeal intakes of the total were 72.0, 12.4, and 8.5 % compared with 71.3, 12.5 and 9.1% for the Tam Hoang and Tau Vang breeds, respectively. The CP intakes as a percentage of total feed intakes were 17.0 and 17.3 % for the Tam Hoang and Tau Vang, respectively, values which were very similar to the balanced feed (17.6 % CP). However, there were significant differences between the mixed and separate diets for daily DM (73.8 and 69.2 g/day) and CP (13.0 and 11.0 g) intakes (P<0.05), but no difference for ME intakes (0.95 and 0.92 MJ/day) (P>0.05).

3.2.2. Egg performance, egg weight and feed consumption per kg eggs

The hen - day production is shown in Table 10, and was significantly higher (P<0.05) for the Tam Hoang (32.7 %) than for the Tau Vang (24.9 %). However, there was no significant difference between the mixed (29.7 %) and separate (27.9 %) diets.

The mean egg weight was significantly different for breed but not significantly different between treatments, and was 44.5 g and 39.5 g for the Tam Hoang and Tau Vang, respectively (P<0.05), and 42.3 and 41.7 g for the mixed and separate diet, respectively.

Feed conversion (FC, kg feed/ kg eggs) was not significantly different between breeds and diets, although the Tam Hoang had somewhat lower FC than the Tau Vang. FC were 5.17 and 5.95 kg feed in DM /kg eggs for the Tam Hoang and Tau Vang, respectively and 5.55 compared with 5.57 kg feed/kg eggs for the mixed and separate diets, respectively. As a result the CP and ME intakes/kg eggs were not significantly different, and were 864g and 67.7 MJ and 1009 g and 77.9 MJ/kg eggs, respectively for the Tam Hoang and Tau Vang hens, and 979g and 71.5 MJ, and 893g and 73.9 MJ/kg eggs for the mixed and separate diets, respectively.

3. Discussion

4.1 Experiment 1

4.1.1 Feed intake

The results for feed intake show that the growing chickens were able to fairly accurately balance their intake of nutrients to meet their requirements, even when the ingredients were supplied separately. This is in agreement with previous studies which have also shown that poultry have a control system that allows them to choose suitable amounts of different foods to satisfy their nutritional requirements, but this is regarded as being too simplistic and other factors such as physical form of ingredients, composition of the food, smell, trough position and previous experience are also likely to be involved (McDonald et al, 1995). Although the amounts of feed and nutrients consumed between the two diets and breeds were not significantly different, the proportion of feedstuffs eaten was different. For example birds on the separate diet consumed more broken rice (84 %),and less soya bean meal (4.7 %) and fishmeal (9.6%) compared with the mixed diet which contained 72 % broken rice, 13 % soya bean meal , and 13 % fishmeal. As result the total CP in the separate diet was only 13.9 % of total DM intake, which is well below the crude protein of the mixed diet (17.1%) and below requirements according to NIAH (1995). Both Tam Hoang and Tau Vang chickens preferred to eat broken rice, which may have been due to an observation that chickens prefer to eat a cereal as grain rather than in the ground form. This result is similar to the findings of Khang (2001) who found that local chickens preferred to eat broken rice rather than soya bean meal. Both Tam Hoang and Tau Vang chickens reduced their intake of soy bean meal to very low levels. This in agreement with Hong Samnang (1997), who found that local chickens ate less protein supplement and more broken rice than an exotic breed, which consumed more protein supplement. However, these differences could also be explained by the fact that the soya bean that was used in the diet was oil extracted soya bean, and the smell was not as attractive as fishmeal

The difference in CP intake thus was considerable, but the two groups still had similar live weight gains. This implies that the protein was balanced and sufficient for the birds' requirement. According to McNaughton et al. (1977 b) the requirements of starting and growing Leghorns at ages of 84 and 140 days are 14 and 20 % CP. Another study (Leeson and Summers, 1979) showed that the estimated requirement for White Leghorn chicks at the age of 0 - 56 days was 12% CP. Requirements for CP for the Vietnamese breeds are likely to be even lower than for the White Leghorn and it seems thus that both the Tam Hoang and Tau Vang were given diets that met or were in excess of their requirements. Amino acid intakes were not different between breeds but were different between diets. This indicates that both improved and local chickens have the ability to balance their nutrient intake, including amino acids, to meet their requirements for growth, and as a result the growth rates of chickens on the separate diet was as high as the group on the mixed diet.

ME intakes were not significantly different between breeds and diets. This indicates that the local breed birds can also adjust their ME intakes to meet their requirement. Even though the proportion of ingredients in the separate diet was different from the mixed diet, and this group ate more broken rice, the difference in total daily ME intake was not significant. This is agreement with the study of Cumming et al. (1987) who showed that each bird can fairly accurately select its intake and balance of nutrients and energy to meet its particular physiological requirements.

4.1.2 Average daily gain

Although the crude protein intake from the two diets was different, the average daily gains of birds on the mixed and separate diets were similar. This indicates that when feed ingredients were supplied separately, chickens can select to meet their requirements and that the CP content of the mixed diet was well in excess of the requirement of both breeds. The Tam Hoang had a higher growth rate than the Tau Vang, because the Tau Vang is a small bodied, unselected local breed while the Tam Hoang is a larger improved dual purpose breed selected for better growth rate and performance. As a result, the efficiency of feed conversion of the Tam Hoang was better than the Tau Vang.

4.1.3 Feed conversion ratio and feed cost per kg gain

Feed cost per kg gain for the Tam Hoang and Tau Vang indicated that the Tau Vang's lower growth rate was the explanation for the higher cost per kg gain. However, the net income for the Tau Vang was not lower than for the Tam Hoang, because consumers, in local markets at least, prefer Tau Vang meat, and the price of a Tau Vang bird is always higher than for the Tam Hoang, normally by about 4,000 to 5,000 VND/kg. The net income for the Tam Hoang on the separate diet was higher than for the Tam Hoang on the mixed diet, but the cost of the mixed diet was higher than for the separate diet. This is due to the fact that the Tam Hoang on the separate diet ate more cheap broken rice and less expensive protein concentrates, but the overall gain was still similar to the gain on the mixed diet, and so the feed cost per kg gain for the separate diet was lower than for the mixed diet. However, even though the Tau Vang on the separate diet also ate large amounts of broken rice the weight gain on the separate diet was lower than on the mixed diet, and so the feed cost per gain was higher than for mixed diet.

4.1.4 Carcass evaluation

The Tam Hoang on the separate diet had high levels of abdominal fat, because they consumed large amounts of broken rice. The extra energy that chickens eat is primarily converted to fat under the skin and abdominal fat (Nguyen Duy Hoan, 1999). This is in agreement with Smith (1990), who reported that heavy breeds will consume more energy on a high energy diet than on a low energy one, and become obese.

4.1.5. Mortality

There was a significant difference in mortality between the Tam Hoang and Tau Vang pullets, which was higher for the Tam Hoang. As the experiment started at a time of changing weather, from dry to rainy season, the chickens, especially the Tam Hoang, which have a poor ability to adapt to hot, humid weather, were very uncomfortable. As a result a number of Tam Hoang died suddenly with no any apparent symptoms. There were no particular diseases observed in this period.

4.2 Experiment 2. Laying period

4.2.1 Feed intake

The local hens consumed about 17 % less feed DM than the improved breed. In contrast with the growing period, when the chickens seemed to prefer fish meal to soya bean meal, in the laying period oil extracted soya bean meal was replaced with roasted soya bean meal, which both the Tam Hoang and Tau Vang obviously liked, as they ate more soya bean than fish meal, and the difference was significant. This indicates that birds offered two kinds of protein eat more of what they like and still can balance nutrient intakes.

The CP intakes were 17.0 and 17.3 % for the Tam Hoang and Tau Vang hens, respectively. These levels are similar the requirement of laying hens according to Smith (2001) who stated that the CP requirement of domestic laying hens in the tropics is 16.5 - 17.5 %. In the laying period hens can adjust the proportion of feed ingredients they consume more accurately than in growing period, As a result nutrient intakes more closely met requirements. However, hens on the mixed and separate diets had significantly different DM and CP intakes, but there was no difference for ME intake, although the egg performance and feed consumption per kg eggs were still not different. This indicates that the diet of the hens on the separate treatment was balanced and still adequate in terms of CP and amino acids. According to Washington (1994) chickens do not require a specific level of crude protein, rather they have a requirement for specific amino acids plus sufficient protein to supply either the non - essential amino acids themselves or amino nitrogen for their synthesis.

4.2.2 Age at first egg and mortality

The Tam Hoang is an improved dual purpose breed. Studies have shown that the breed has late sexual maturity, especially the Tam Hoang 882 strain, which lays its first egg at 161 days according to Le Hong Son (1997). However, in our study the Tam Hoang birds were of the Luong Phuong strain. In addition, they were reared on station in early summer when day length was increasing, which could be the reason that the first egg of the Tam Hoang in our experiment was as early as 127 days. According to Rose (1997) the mean age that a flock of poultry reaches sexual maturity depends upon species and strain, but changing day lengths during rearing also have a large influence. The Tau Vang is local dual purpose breed, rather than a layer breed, which could explain why the first egg of Tau Vang was10 days later than the Tam Hoang.

The mortality rates for the Tam Hoang and Tau Vang hens were 15.0 % and 29.8 %, respectively (P<0.05). The laying period was from July to November, so the Tam Hoang in this period had adapted to the hot weather. Tau Vang deaths were caused by Mareks disease, against which they were not vaccinated. However, the Tam Hoang, purchased in Ho Chi Minh City, were vaccinated against Mareks disease.

4.2.3 Egg performance, egg weight and feed consumption per kg eggs

The Tam Hoang's egg production was higher than the Tau Vang's at 32.7 % and 24.9 % respectively, for the 20 weeks of laying. The highest egg rate was from 8 to 13 weeks, and after that egg performance decreased slowly, which is the usual pattern. As the Tam Hoang reached sexual maturity earlier than the Tau Vang, so the highest egg rate was between week 8 to 13, while for the Tau Vang it was between week 11 to 14 after the first egg. A similar difference in egg performance between local and improved breeds was found by Do Viet Minh (1999), who reported that the hen-day production from Tam Hoang and Ri (an unimproved local breed common in North Vietnam) hens was 39.5 and 17.9 % for 14 weeks of laying, which also shows that the Ri hen has lower egg performance than the Tau Vang and Tam Hoang breeds.

4.2.4 Mean egg weight

The egg weight for the on-station hens was slightly higher than for the same breed reared on farm (Paper II). This agrees with a study by Do Viet Minh (1999), who showed that egg weight is affected by nutrient intake, and also depends on breed and strain, and in particular on mature body weight. The egg weights were nearly similar on the two diets, however, which indicates that the different nutrient and ME intakes from the mixed and separate diets did not affect egg weight.

4.2.5 Economic analysis

The feed costs per kg eggs for the two breeds were not significantly different, although the feed cost per kg of Tam Hoang eggs was slightly lower than for the Tau Vang (15,302 and 17,6652 VND/kg eggs, respectively (P>0.05), which is 13.7 % lower for the improved hens compared with local hens. This difference is due to the lower egg performance of the Tau Vang. However, the feed consumption for 1 egg was similar (230 and 235 g/egg for the Tan Hoang and Tau Vang, respectively) (P>0.05). Consequently, as market prices per egg are similar for the two breeds, raising Tam Hoang and Tau Vang hens give similar economic benefits, at least under on-station conditions. The results from this study are quiet different from the results of Do Viet Minh (1999), who found that the feed consumption per kg eggs for the Tam Hoang was lower than for the Ri by about 36 %.

There was no significant difference between the two treatments in feed costs per kg eggs and also CP and ME required to produce 1 kg eggs were not significantly different. It seems that both Tam Hoang and Tau Vang balance their feed and nutrient intakes for maximum production and as a result the economic benefits on the mixed and separate diets were similar

5. Conclusions

DM, CP and ME intakes for the two breeds in the growing period were not different, but were in the laying period. However, intakes of CP and amino acids were lower on the separate diet than on the mixed diet. Average daily gain, egg performance and feed consumption per kg eggs were not different between the mixed and separate diets.

The growth rate of the Tam Hoang was higher than of the Tau Vang, but the feed conversion ratio was not different between the two breeds.

The feed consumption of the Tam Hoang hens was 18 % higher than of the local Tau Vang breed, and also egg performance of the Tam Hoang was higher than the Tau Vang. However, feed conversion per kg eggs was not different between treatments and breeds, which indicates that both improved and local breed hens seem able to balance their nutrient intakes to meet their requirements for body maintenance and egg production.

Acknowledgements

Appreciation is expressed to the Swedish Agency for Research Co-operation with Developing Countries (SAREC) and the Swedish University of Agricultural Sciences (SLU) for its financial and material support of this study.

I am also indebted to the Department of Animal Husbandry, College of Agriculture and Experimental farm for allowing me to use the facilities and chicken house for my work.

I would like to thank to my supervisor, Dr. Brian Ogle who has spent a lot of time to correct and guide my studies. Many thanks to my teacher Mrs.La Thi Thu Minh for her help and devoted guidance, to the head of the experimental farm for his enthusiastic help and encouragement. Also thanks to the staff of the experimental farm for their help and cooperation.

Many thanks to my colleagues Kim Khang and my assistants Hau, Quyen, Thoai, Giau, Long, Linh, Binh, Ut, Quan for their valuable help.

Special thanks to my family for their encouragement and help, to my husband Hong Tien for his help and love, to my daughter Hong Nhung for her love.

References

AOAC.1980. Official Methods of Analysis, Association of official Analytical Chemists, 13th Ed.

Cumming, R. B., Mstika, I. M.,Wodzicka, T.M., 1987. Practical aspects of choice feeding in poultry and its future role, CAB abstracts 1987 -1989.

Dessie, T.1996. Studies on village poultry production systems in the central highlands of Ethiopia, Msc. Thesis, Department of Animal Nutrition and Management, Swedish University of Agricultural Sciences (SLU), Uppsala, 1996.

Gunaratne, S.P., Charndrasiri, A.D., Mangalika Hemalatha, W.A.P., Roberts, J.A. 1992. The productivity and nutrition of Village Chickens in Srilanka.

Hong Samnang. 1997. Effect of breed (Local versus exotic hybrids) on selection of dietary ingredients in a scavenging system for growing hens. The report on feed resources in sustainable livestocked - based Agriculturre in SE Asia, Workshop 1997.

Hoan, N. D. 1999. The effects of energy levels on the low protein diets on the abdominal organs, lipit metabolisable and abdominal fat of laying hens. Special Issue, Vietnam Animal husbandry society, Hanoi, 1999.

Janssen, W. M. A., ed. 1989. European Table of Energy Values for poultry feedstuffs. 3rd ed. Beekbergen, Netherlands: Spelderholt Center for Poultry Research and Information Services.

Khang, N.T.K. 2001. Scavenging behavior and feed selection by exotic and local chickens, Miniproject, UTA Foundation, Chamcar Daung, Dangkor District, PO Box 2423, Phnom Penh 3, Cambodia.

Leeson, S. and Summers, J.D. 1979. Step up protein diets for growing pullets. Poult. Sci.58:681

Minh, D.V.1999. Effects of energy and protein supplementation strategy on production performance of local and improved scavenging hens in North Vietnam. Msc.Thesis Swedish University of Agricultural Sciences, Uppsala, Sweden.

McDonald. P., Edwards, R.A., Greenhalgh, J. F.D., Morgan, C.A. 1995. Animal nutrition, fifth Edition, Longman Scientific and Technical, copublished in the United States with John Wiley and Sons, Inc. New York. 418 -424

NIAH (National Institute of Animal Husbandry). 1995. Composition and nutritive value of animal feeds in Vietnam. Agriculture Publishing House. Hanoi, Vietnam. 251 pp.

Rose, S.P.1997. Priciples of poultry Sciences, CAB International, London

Smith, A.J. 1990. Poultry tropical agriculturist series.CTA, Macmillan publishers, London. 47, 55 - 71, 179. And second edition 2001.

Son, L. H and Tien, H.V. 1997.Scientific papers on animal production, 1996 -1997, Agricultural Publishing House, Hanoi -1998, 54 -65, 66 -79

Van Soest, P.J., Mason, V.C.1991. The influence of the Maillard reaction upon the nutritive value of fibrous feeds. Anim. Feed sci.Tech.32:45-53.

Washington, D.C. 1994. Nutient requirements of Poultry, ninth revised edition, 1994. 9 -33, 85-90.


Table 1. Chemical composition of feedstuffs and the mixed diet, growing period (in DM)

Item

Broken rice

Soya bean meal

Fish meal

Mixed diet

DM (%)

87.5

89.0

89.0

87.8

CP (%)

8.3

41.0

48.9

17.1

EE (%)

0.6

3.4

3.0

1.2

CF (%)

0.6

4.4

3.6

1.5

Ash (%)

1.2

3.8

29.9

5.0

NFE (%)

76.8

39.0

4.0

61.5

NDF (%)

1.3

8.7

2.4

2.2

Ca (%)

1.3

0.9

5.3

0.97

P (%)

0.28

0.7

2.5

0.72

Lysine (g/kg)

3.2

16.5

30

8.5

Methionine,g/kg

1.8

5.3

14.4

4.0

ME (MJ/kg)

14.41

9.7

9.8

12.9

(Calculated)

Table 2. Essential amino acid composition of feed ingredients and the mixed diet, growing period

Amino acid

% in DM

Broken rice

Soya bean meal

Fish meal

Mixed diet

Lysine

0.32

1.66

3.0

0.84

Isoleucine

0.32

1.71

2.09

0.73

Leucine

0.65

2.93

3.49

1.31

Methionine

0.19

0.53

1.44

0.39

Threonine

0.23

1.46

2.15

0.64

Valine

0.44

1.9

2.45

0.89

Phenylalanine

0.34

2.0

1.98

0.77

Histidine

0.28

2.05

1.72

0.69

Arginine

0.57

2.62

4.09

1.29

Tryptophan

0.12

0.41

1.2

0.28

Table 3. Effect of breed on DM and nutrient intake, FCR and average daily gain, growing period

Item

Breed

SE

P-value

TH

TV

Initial weight, g

609

319

71.53

0.000

Final weight, g

2130

1290

90

0.001

ADG, g *

20.5

10.7

1.45

0.009

Total DM intake, g/day *

61.2

40.1

5.76

0.073

CP intake, g/day *

9.14

6.33

1.01

0.061

CP intake,% of total DM

15.5

16.5

ME intake, MJ/day *

0.72

0.47

0.07

0.092

FCR, kg/kg *

2.95

3.61

0.40

0.716

CP/gain (g/kg) *

430

590

84

0.364

ME/gain, MJ/kg *

34.9

43.1

5.01

0.426

CP/ME, g/MJ *

12.74

13.7

0.7

0.505

Lysine,g

0.42

0.32

0.05

0.307

Methionine, g

0.21

0.14

0.02

0.174

Threonine

0.32

0.24

0.04

0.326

Mortality, %

13.4

5

2.42

0.029

(*) Adjusted by covariance according to initial weight

Table 4. Effect of dietary treatment on DM and nutrient intakes, FCR and ADG

Items

Diet

SE

P-value

Mixed

Separate

Initial weight, g

464

464

0.007

0.971

Final weight, g

1740

1670

15

0.013

Total DM intake, g/day *

49.9

51.4

0.96

0.309

CP intake, g *

8.51

6.95

0.17

0.000

CP intake,% of total DM

17.1

13.9

ME intake, MJ/day *

0.58

0.61

0.012

0.074

FCR *

3.15

3.41

0.068

0.018

ADG, g *

15.9

15.3

0.24

0.157

CP/gain (g/kg) *

550

480

14

0.006

ME/gain (MJ/kg) *

37

40.9

0.84

0.007

CP/ME (g/MJ) *

14.81

11.63

0.11

0.000

Lysine,g

0.42

0.32

0.008

0.000

Methionine, g

0.19

0.16

0.004

0.000

Threonine

0.32

0.24

0.006

0.000

Mortality (%)

9.8

8.6

2.42

0.716

(*) Adjusted by covariance according to initial weight

Table 5. Effect of treatment and breed on the proportion of the ingredients of total daily feed intake, growing period (%).

Item

Treatment

Breed

Mixed

Sep

SE

P-value

TH

TV

SE

P-value

Broken rice, %

72.0

84.0

0.40

0.00

79.3

76.6

0.40

0.000

Soya bean, %

13.0

4.7

0.42

0.00

8.3

9.4

0.42

0.073

Fish meal,%

13.0

9.6

0.15

0.00

10.5

12.1

0.15

0.000

Oyster, bone meal, %

2.0

1.7

0.01

1.9

1.8

CP %

17.1

13.9

15.5

16.5

Table 6. Effect of breed and treatment on relative intakes of the dietary ingredients (%)

Item

Tam Hoang

Tau Vang

SE

P-value

Mixed

Sep.

Mixed

Sep.

Broken rice,%

72.0

86.6

72.0

81.3

0.56

0.000

Soya bean,%

13.0

3.5

13.0

5.8

0.60

0.000

Fish meal,%

13.0

8.1

13.0

11.2

0.21

0.000

Oyster, bone meal, %

2.0

1.7

2.0

1.7

0.01

CP, %

17.1

12.9

17.1

14.9


Table 7. Effect of breed and dietary treatment on carcass characteristics.

Iterm

Breed (B)

Diet (D)

P-value

Tam

Tau Vang

Mixed

Sep

Breed (B)

Diet (D)

B*D

Hoang

Live weight, g

1989

1178

1603

1564

0.00

0.16

0.07

Carcass weight, g *

1162

997

1097

1062

0.23

0.04

0.96

% carcass

72

63

69

66

0.24

0.03

0.62

Breast muscle, g *

253

236

253

236

0.83

0.04

0.17

Thigh muscle,g *

199

179

197

180

0.58

0.00

0.12

Gizzard /carcass ,%

3.6

6.0

4.7

5.0

0.53

0.61

0.40

Abdominal fat, g *

99.4

29.1

43.1

85.3

0.02

0.00

0.14

Abd fat/carcass, %

7.2

3.7

4.0

7.0

0.06

0.01

0.22

Liver/carcass %

4.5

2.4

3.4

3.5

0.12

0.80

0.05

(*)Adjusted by covariance according to live weight

2. Laying period

Table 8. Effects of breed on egg production, egg weight, feed conversion and mortality.

Items

Breed

SE

P-value

Tam Hoang

Tau Vang

Daily feed intake (g) (*)

78.2

64.9

7.1

0.364

Daily CP intake (g) (*)

13.2

10.8

1.3

0.474

CP intake,% of total DM

17.0

17.3

Daily ME intake (MJ) (*)

1.02

0.84

0.1

0.363

Total eggs

3095

1865

Hen- day production (%)

32.7

24.9

1.93

0.015

Egg weight (g)

44.5

39.5

0.47

0.000

FCR (kg feed/kg eggs)

5.17

5.95

0.34

0.135

CP/kg eggs (g/kg )

864

1009

60

0.112

ME/kg eggs (MJ/kg)

67.6

77.9

4.5

0.135

Feed cost/kg eggs (VND/kg)

15,302

17,655

1044

0.137

Mortality (%)

15.0

29.8

4.6

0.044

AWG (g/day)

3.8

3.9

0.17

0.686

(*) Adjusted by covariance according to initial weight


Table 9.Effect of dietary treatment on egg production, egg weight, feed per kg eggs and mortality.

Items

Treatment

SE

P-value

Mixed

Separate

Daily feed intake (g) (*)

73.8

69.2

1.20

0.023

Daily CP intake (g) (*)

13.0

11.0

0.23

0.000

CP intake,% of total DM

17.6

16.7

Daily ME intake (MJ) (*)

0.95

0.92

0.02

0.205

Total eggs

2456

2504

Hen-day production (%)

29.68

27.96

1.93

0.541

Egg weight (g/)

42.31

41.71

0.47

0.390

FCR/ (kg feed/kg eggs)

5.55

5.57

0.34

0.960

CP/kg eggs (g/kg )

979

894

60

0.331

ME/kg eggs (MJ/kg)

71.5

73.9

4.5

0.713

Feed cost /kg eggs (VND/kg)

16,756

16,202

1044

0.714

Mortality (%)

27.8

17.2

4.6

0.134

AWG (g/day)

4.2

3.5

0.17

0.024

(*) Adjusted by covariance according to initial weight

Table 10. Effect of breed and treatment on egg production, egg weight and feed conversion

Item

Tam Hoang

Tau Vang

SE

P-value

Mixed

Sep.

Mixed

Sep.

Daily FI (g) (*)

81.8

74.5

65.8

63.9

7.20

0.156

Daily CP I (g) (*)

14.3

11.6

11.6

10.5

1.37

0.032

CP ,% of total DM

17.6

16.4

17.6

16.9

Daily ME I (MJ) (*)

1.06

1

0.84

0.84

0.1

0.228

Total eggs

1531

1564

925

940

Hen- day, (%)

33.2

32.3

26.27

23.6

2.7

0.742

Egg weight (g)

44.9

44.1

39.6

39.3

0.67

0.690

FCR/ (kg /kg egg)

5.42

4.92

5.67

6.22

0.48

0.299

CP/kg egg (g/kg )

956

772

1003

1015

84.8

0.272

ME/kg egg (MJ/kg)

69.8

65.4

73.3

82.4

6.4

0.310

Feed cost/kg egg

(VND/kg)

16,346

14,258

17,166

18,145

1477

0.320

Mortality (%)

20.0

10.0

35.0

24.0

6.6

0.926

AWG (g/day/hen)

4.2

3.5

4.27

3.6

0.24

0.919

(*) Adjusted by covariance according to initial weight


Table 11. Effect of dietary treatment on daily feed and ingredient intake (%)

Items

Dietary treatment

Breed

Mix

Sep

SE

P-value

TH

TV

SE

P-value

MM, %

71.0

72.4

0.15

0.000

72.0

71.3

0.15

0.006

SBM, %

10.0

14.9

0.18

0.000

12.4

12.5

0.18

0.638

FM, %

12.0

5.6

0.06

0.000

8.5

9.1

0.06

0.000

Oyster, BM,%

7.0

7.0

7.0

7.0

CP%

17.6

16.7

17.0

17.3

Table 12. Effect of breed and treatment on relative intakes of the dietary ingredients (%).

Item

Tam Hoang

Tau Vang

SE

P-value

Mixed

Sep.

Mixed

Sep.

Maize meal, %

71.0

73.1

71.0

71.6

0.21

0.006

Soya bean meal, %

10.0

14.8

10.0

15.0

0.25

0.000

Fish meal, %

12.0

5.0

12.0

6.2

0.09

0.000

Oyster, bone meal,%

7.0

7.0

7.0

7.0

CP, %

17.6

16.4

17.6

16.9